Professor Andrew Loveridge
Deputy Director
Kaplan Research Fellow Lady Margaret Hall
Andrew Loveridge is the Kaplan Senior Research Fellow at Lady Margaret Hall and the Wildlife Conservation Unit, Oxford University. Born and raised in Zimbabwe, he read for his doctoral degree in Zoology at Oxford, and after graduating embarked on a career as an ecologist and conservationist. Since 1999 he has run a long-term African lion research and conservation project, initially based in hwange National Park, Zimbabwe, but now covering much of the southern part of the Kavango-Zambezi Transfrontier Conservation Area, under the banner of the Trans-Kalahari Predator Programme (TKPP). Over the last twenty years his research and conservation work on lions has encompassed lion population management, impacts of trophy hunting, socio-biology, behavior, predator-prey relationships, human-wildlife conflict and its mitigation landscape ecology, and trade. He has published over 100 peer reviewed papers and book chapters. In 2012 he received the SATIB Trust Award for contributions to African Conservation. He is a member of the IUCN Cat Specialist Group, Canid Specialist Group and African Lion Working Group and the KAZA Carnivore Conservation Coalition.
Loveridge is an experienced editor and author, having co-edited Biology and Conservation of Wild Felids (OUP, 2010) and he serves as an Associate Editor for African Journal of Wildlife Research. He recently published an auto-biographical account of his research and conservation work, Lion-hearted (Regan Arts, New York 2018), which was included in the Forbes Ten of the Best Books About Climate Change, Conservation, and The Environment of 2018.
Selected Publications
Barnier, F., M. Valiex, P. Duncan, S. Chamaillé-Jammes, P. Barre, A. J. Loveridge, D. W. Macdonald, and H. Fritz. 2014. Diet quality in a wild grazer declines under the threat of predation. Proceedings of the Royal Society B 281.
Benhamou, S., M. Valeix, S. Chamaillé-Jammes, D. W. Macdonald, and A. J. Loveridge. 2014. Movement-based analysis of interactions in African lions. Animal Behaviour http://dx.doi.org/10.1016/j.anbehav.2014.01.030.
Brown, C.R., Pienke, D., & Loveridge, A. J. (1996). Mortality in near-term ostrich embryos during artificial incubation. British Poultry Science, 37, 73-85.
Crosmary, W., Loveridge, A. J., Ndaimani, H., Lebel, S., Booth, V., Cote, S., & Fritz, H. (In Press). Trophy hunting in Africa: long-term trends in antelope horn size. Animal Conservation. doi: 10.1111/acv.12043
Davidson, Z., Valeix, M., Loveridge, A.J., Hunt, J. E., Johnson, P.J., Madzikanda, H., & Macdonald, D.W. (2012). Environmental determinants of habitat and kill site selection in a large carnivore: scale matters. Journal of Mammalogy, 93(3).
Davidson, Z., Valeix, M., Loveridge, A.J., Madzikanda, H., & Macdonald, D.W. (2011). Socio-spatial behaviour of an African lion population following perturbation by sport hunting. Biological Conservation, 144, 114-121.
Davidson, Z., Valeix, M., van Kesteren, F., Loveridge, A. J., Hunt, J. E., Murindagomo, F., & Macdonald, D. W. (2013). Seasonal diet and prey preference of the African lion in a waterhole-driven semi-arid savanna. PLoS ONE, 8(2), 1-10.
Davidson, Z., Valeix, M., van Kesteren, F., Loveridge, A. J., Hunt, J. E., Murindagomo, F., & Macdonald, D. W. (in press). Seasonal diet and prey preference of the African lion in a waterhole-driven semi-arid savanna. PLoS ONE.
duPreez, B., T. Hart, A. J. Loveridge, and D. W. Macdonald. 2015. The impact of risk on animal behaviour and habitat transition probabilities. Animal Behaviour 100:22-37.
duPreez, B., A. J. Loveridge, and D. W. Macdonald. 2014a. Making the best of camera-trap surves in an imperfect world. Biological Conservation DOI 10.1016/j.biocon.2014.09.022.
duPreez, B., A. J. Loveridge, and D. W. Macdonald. 2014b. To bait or not to bait: a comparison of two camera-trapping methods for estimating leopard Panthera pardus density. Biological Conservation 176 153–161.
Elliot, N., S. A. Cushman, D. W. Macdonald, and A. J. Loveridge. 2014a. The devil is in the dispersers. Predictions of landscape connectivity change with demography. Journal of Applied Ecology 51:1169-1178.
Elliot, N., D. W. Macdonald, A. J. Loveridge, and S. A. Cushman. 2014b. Movements vary according to dispersal status, group size and rainfall: the case of the African lion. Ecology 95:2860-2869.
Elliot, N., M. Valiex, D. W. Macdonald, and A. J. Loveridge. 2014c. Social relationships affect dispersal timing revealing a delayed infanticide in African lions. Oikos 123:1049-1056.
Fahlman, A., Loveridge, A., Wenham, C., Foggin, C., Arnemo, J. M., & Nyman, G. (2005). Reversible anaesthesia of free-ranging lions (Panthera leo) in Zimbabwe. Journal of the South African Veterinary Association., 76, 187-192.
Funston, P.J., Frank, L., Stephens, T., Davidson, Z., Loveridge, A.J., Macdonald, D.W., . . . Ferreira, S. (2010). Substrate and species constraints on the use of track incidences to estimate African large carnivore abundance. Journal of Zoology, 281, 56-65.
Hodgeson, A. N., Hanel, C., & Loveridge, A. J. (1994). The reproductive cycle of the pulmonate snail Xerocerastes minutus from the Namib desert. Journal of Molluscan Studies, 60, 355-358.
Hunter, L.T.B., White, P.A., Henschel, P.P., Frank, L., Burton, A.C., Loveridge, A. J., . . . Breitenmoser, U. (2013a). No science, no success and still no need for captive- origin lion reintroduction: a reply to Abell and Youldon. Oryx, 47(1), 27-28.
Hunter, L.T.B., White, P.A., Henschel, P.P., Frank, L., Burton, C., Loveridge, A. J., . . . Breitenmoser, U. (2013b). Walking with lions: Why there is no role for captive origin lions (Panthera leo) in species restoration. Oryx, 14(1), 19-24.
Johnson, P.J., Kansky, R., Loveridge, A.J., & Macdonald, D.W. (2010). Size, Rarity and Charisma: Valuing African Wildlife Trophies. PLoS ONE, 5(9). e12866
Kesch, K. M., D. T. Bauer, and A. J. Loveridge. 2014. Undermining game fences: who is digging holes in Kalahari sands. African Journal of Ecology online early.
Loveridge, A. J. , & Macdonald, D. W. (2002). Habitat ecology of two sympatric species of jackals in Zimbabwe. Journal of Mammalogy, 83(2), 599-607.
Loveridge, A. J., & Macdonald, D.W. (2003). Niche separation in sympatric jackals (Canis mesomelas and Canis adustus). Journal of Zoology, London, 259, 143-153.
Loveridge, A.J., Hunt, J.E., Murindagomo, F., & Macdonald, D.W. (2006). Influence of drought on predation of elephant (Loxodonta africana) calves by lions (Panthera leo) in an African wooded savannah. Journal of Zoology, 270(523-530).
Loveridge, A.J., Searle, A.W., Murindagomo, F., & Macdonald, D.W. (2007). The impact of sport hunting on the population dynamics of an African lion population in a protected area. Biological Conservation, 134, 548-558.
Loveridge, A.J., Valeix, M., Davidson, Z., Murindagomo, F., Fritz, H., & Macdonald, D.W. (2009). Changes in home range size of African lions in relation to pride size and prey biomass in a semi-arid savanna. Ecography, 32, 1-10.
Loveridge, Andrew. J., & Macdonald, David. W. (2001). Seasonality in spatial organization and dispersal of sympatric jackals (Canis mesomelas and C. adustus): implications for rabies management. Journal of Zoology, 253, 101-111.
Loveridge, A. J., & Macdonald, D. W. (2013). Canis adustus- Side-striped jackal. In J. Kingdon & M. Hoffmann (Eds.), Mammals of Africa (Vol. V). London: Bloomsbury Books.
Loveridge, A. J., & Nel, J.A.J. (2013). Canis mesomelas- Black-backed jackal. In J. Kingdon & M. Hoffmann (Eds.), Mammals of Africa (Vol. V). London: Bloomsbury Books.
Packer, C., Loveridge, A. J., Canney, S., Caro, T., Garnett, S.T., Pfeifer, M.L., . . . Polasky, S. (2013). Conserving Large Carnivores: Dollars and Fence Ecology Letters, 16(5), 635-641. doi: DOI:10.1111/ele.12091
Packer, C., Swanson, A., Canney, S., Loveridge, A. J., Garnett, S.T., Pfeifer, M.L., . . . MacNulty, D. (In Press). The case for fencing remains intact. Ecology Letters, DOI:10.1111/ele.12171. doi: doi: 10.1111/ele.12171
Periquet, S., Todd-Jones, L., Valeix, M., Stapelkamp, B., Elliot, N., Wijers, M., . . . Loveridge, A. J. (2012). Influence of immediate predation risk by lions on the vigilance of prey of different body size. Behavioral Ecology.
Periquet, S., Valeix, M., Loveridge, A.J., Madzikanda, H., Macdonald, D.W., & Fritz, H. (2010). Individual vigilance of African herbivores while drinking: the role of immediate predation risk and context. Animal Behaviour, 79, 665-671.
Pfeifer, M. L., C. Packer, A. C. Burton, S. T. Garnett, A. J. Loveridge, D. MacNulty, and P. Platts. 2014. In defence of fences: response to Woodroffe et al. Science 345:389.
Prescott, G., Johnson, P.J., Loveridge, A.J., & Macdonald, D.W. (2012). Does change in IUCN status affect demand for African bovid trophies? Animal Conservation, 15, 248-252. doi: doi:10.1111/j.1469-1795.2011.00506.x
Valeix, M., Chamaillé-Jammes, S., Loveridge, A.J., Davidson, Z., E., Hunt. J., Madzikanda, H., & Macdonald, D.W. (2011). Understanding patch departure rules for large carnivores: Lion movements support a patch disturbance hypothesis. The American Naturalist, 178(2), DOI: 10.1086/660824.
Valeix, M., Fritz, H., Loveridge, A.J., Davidson, Z., Hunt, J. E., Murindagomo, F., & Macdonald, D.W. (2009). Does the risk of encountering lions influence African herbivore behaviour at waterholes? Behavioural Ecology and Sociobiology, 63, 1483-1494.
Valeix, M., Hemson, G., Loveridge, A.J., Mills, G., & Macdonald, D.W. (2012). Behavioural adjustments of a large carnivore to access secondary prey in a human dominated landscape. Journal of Applied Ecology, 49, 73-81.
Valeix, M., Loveridge, A. J., & Macdonald, D. W. (2012). Influence of prey dispersion and group size of African lions: a test of the resource dispersion hypothesis. Ecology, 93(11), 2490–2496.
Valeix, M., Loveridge, A.J., Chamaillé-Jammes, S., Davidson, Z., Murindagomo, F., Fritz, H., & Macdonald, D.W. (2009). Behavioural adjustments of African herbivores to predation risk by lions: Spatiotemporal variations influence habitat use. Ecology, 90(1), 23-30.
Valeix, M., Loveridge, A.J., Davidson, Z., Madzikanda, H., Fritz, H., & Macdonald, D.W. (2010). How key habitat features influence large terrestrial carnivore movements: waterholes and African lions in a semi arid savanna of north-western Zimbabwe. Landscape Ecology, 25, 337-351.
Book chapters and published reports
Atkinson, R. P. D., & Loveridge, A. J. (2004). Side-striped jackal, Canis adustus. In Claudio Sillero-Zubiri, Michael Hoffman & David W. Macdonald (Eds.), Canids: Foxes, Wolves, Jackals and Dogs. Status Survey and Conservation Action Plan (pp. 152-156). Gland, Switzerland and Cambridge, UK: IUCN, SSC Canid Specialist Group.
Loveridge, A. J. (2001). Jackals. In D. W. Macdonald (Ed.), The New Encyclopaedia of Mammals (pp. 50-51). Oxford Oxford University Press.
Loveridge, A. J., Lynam, T., and Macdonald, D.W. (2006). Lion Conservation Research. Workshops 3 & 4: From conflict to Socioecology (pp. 95).
Loveridge, A. J., & Nel, J. A. J. (2004). Black-backed jackal, Canis mesomelas. In Claudio Sillero-Zubiri, Michael Hoffman & David W. Macdonald (Eds.), Canids: Foxes, Wolves, Jackals and Dogs. Status Survey and Conservation Action Plan (pp. 161-166). Gland, Switzerland and Cambridge, UK: IUCN, SSC Canid Specialist Group.
Loveridge, A. J., Reynolds, J. C., & Milner-Gulland, E. J. . (2007). Does sport hunting benefit conservation? In D. W. Macdonald & K. Service (Eds.), Key Topics in Conservation (pp. 224-240). Oxford, UK: Blackwell Publishing.
Loveridge, A.J., & Canney, S. (2009). Report on the lion distribution and conservation modelling project: Report to the Born Free Foundation, UK.
Loveridge, A.J., Hemson, G., Davidson, Z., & Macdonald, D.W. (2010). African lions on the edge: Reserve boundaries as ‘attractive sinks’. In D.W. Macdonald & A.J. Loveridge (Eds.), Biology and Conservation of Wild Felids (pp. 283- 304). Oxford: Oxford University Press.
Loveridge, A.J., Packer, C., & Dutton, A. (2009). Science and the recreational hunting of lions. In B. Dickson, J. Hutton & W.M. Adams (Eds.), Recreational hunting, conservation and rural livelihoods (pp. 108-124). Oxford: Wiley-Blackwell.
Loveridge, A.J., Wang, S.W., Frank, L. G., & Seidensticker, J. (2010). People and wild felids: conservation of cats and management of conflicts. In D.W. Macdonald & A.J. Loveridge (Eds.), Biology and Conservation of Wild Felids (pp. 161-196). Oxford: Oxford University Press.
Loveridge, Andrew J., Lynam, Tim, & Macdonald, David W. (Eds.). (2001). Lion Conservation Research. Workshop 1: Survey Techniques. Oxford, UK: Wildlife Conservation Research Unit, University of Oxford.
Loveridge, Andrew. J., Lynam, A., & Macdonald, D.W. (2002). Lion Conservation Research Workshop 2: Modelling Conflict. Oxford: WildCRU and Darwin Initiative.
Loveridge, A. J., & Macdonald, D. W. (2013). Canis adustus- Side-striped jackal. In J. Kingdon & M. Hoffmann (Eds.), Mammals of Africa (Vol. V). London: Bloomsbury Books.
Loveridge, A. J., & Nel, J.A.J. (2013). Canis mesomelas- Black-backed jackal. In J. Kingdon & M. Hoffmann (Eds.), Mammals of Africa (Vol. V). London: Bloomsbury Books.
Macdonald, D.W., Loveridge, A.J., & Atkinson, R.P.D. (2004). A comparative study of side-striped jackals Canis adustus in Zimbabwe: the influence of habitat and congeners. In D.W. Macdonald & C. Sillero-Zubiri (Eds.), The biology and conservation of wild canids. (pp. 255-270). Oxford, UK: Oxford University Press.
Macdonald, D.W., Loveridge, A.J., & Nowell, K. (2010). Dramatis personae: An introduction to the wild felids. In D.W. Macdonald & A.J. Loveridge (Eds.), Biology and Conservation of Wild Felids (pp. 3-61). Oxford: Oxford University Press.
Macdonald, D.W., Loveridge, A.J., & Rabinowitz, A. (2010). Felid futures: crossing disciplines, borders, and generations. In D.W. Macdonald & A.J. Loveridge (Eds.), Biology and Conservation of Wild Felids (pp. 599-653). Oxford: Oxford University Press.
Packer, C., Whitman, K., Loveridge, A., Jackson, J., & Funston, P. (2006). Impacts of trophy hunting on lions in East and Southern Africa: Recent offtake and future recommendations. Paper presented at the The Eastern and Southern African Lion conservation Workshop, Johannesburg, South Africa.